Pdf's of most publications can be accessed through my profile:


22. Nanglu, K., Cole, S.R., Wright, D.F., and Souto, C. Submitted. Worms and gills, plates and spines: the evolutionary origins and incredible disparity of deuterostomes revealed by fossils, genes, and development. Biological Reviews.

21. Cole, S.R., and D.F. Wright. Accepted. Niche evolution and phylogenetic community paleoecology of Late Ordovician crinoids. Elements of Paleontology.


20. Cole, S.R. 2022. Early echinoderms decouple form and function. Nature Ecology & Evolution. [pdf]

19. Fawcett, S., D. Agosti, S.R. Cole, and D.F. Wright. 2022. Digital accessible knowledge: mobilizing legacy data and the future of taxonomic publishing. Bulletin of the Society of Systematic Biologists.

18. Cole, S.R., and M.J. Hopkins. 2021. Selectivity and the effect of mass extinctions on disparity and functional ecology. Science Advances.

17. Cole, S.R., W.I. Ausich, and M.A. Wilson. 2020. A Hirnantian holdover from the Late Ordovician mass extinction: phylogeny and biogeography of a new anthracocrinid crinoid from Estonia. Papers in Palaeontology.

16. Cole, S.R., D.F. Wright, W.I. Ausich, and J.M. Koniecki. 2020. Paleocommunity composition, relative abundance, and new camerate crinoids from the Brechin Lagerstätte (Upper Ordovician). Journal of Paleontology.

15. Zamora, S., D.F. Wright, R. Mooi, B. Lefebvre, T.E. Guensburg, P. Gorzelak, B. Da14. vid, C.D. Sumrall, S.R. Cole, A. Hunter, J. Sprinkle, J.R. Thompson, T. Ewin, O. Fatka, E. Nardin, M. Reich, M. Nohejlová, and I.A. Rahman. 2020. Re-evaluating the phylogenetic position of the enigmatic early Cambrian deuterostome Yanjiahella. Nature Communications.

14. Ausich, W.I., D.F. Wright, S.R. Cole, and G. Sevastopulo. 2020. Homology of posterior interray plates in crinoids: a review and new perspectives from phylogenetics, the fossil record, and development. Palaeontology.

​​13. Wright, D.F., S.R. Cole, and W.I. Ausich. 2019. Biodiversity, systematics, and new taxa of cladid crinoids from the Ordovician Brechin Lagerstätte. Journal of Paleontology.

12. Cole, S.R. 2019. Hierarchical controls on extinction selectivity across the diplobathrid crinoid phylogeny. Paleobiology.

11. Cole, S.R., D.F. Wright, and W.I. Ausich. 2019. Phylogenetic community paleoecology of one of the earliest complex crinoid faunas (Brechin Laerstätte, Ordovician). Palaeogeography, Palaeoclimatology, Palaeoecology, v. 521, p. 82-98.
10. Cole, S.R. 2018. Phylogeny and evolutionary history of diplobathrid crinoids (Echinodermata). Palaeontology, v. 91, p. 815-828.

9. Ausich, W.I., D.F. Wright, S.R. Cole, and J. Koniecki. 2018. Disparid and hybocrinid crinoids (Echinodermata) from the Upper Ordovician (lower Katian) Brechin Lagerstätte of Ontario. Journal of Paleontology, v. 92, p. 850-871.

8. Cole, S.R., and U. Toom. 2018. Two new camerate crinoid genera from the Upper Ordovician (Katian) of Estonia: evolutionary origin of Family Opsiocrinidae and a phylogenetic assessment of Ordovician Monobathrida. Journal of Systematic Paleontology. 519

7. Cole, S.R, W.I. Ausich, D.F. Wright, and J. Koniecki. 2018. An echinoderm Lagerstätte from the Upper Ordovician (Katian), Ontario: taxonomic re-evaluation and description of new dicyclic camerate crinoids. Journal of Paleontology, v. 92, p. 488-505.

6. Cole, S.R. 2017. Phylogeny and morphologic evolution of the Ordovician Camerata (Class Crinoidea, Phylum Echinodermata). Journal of Paleontology, v. 91, p. 815–828.

5. Cole, S.R., W.I. Ausich, S. Zamora, and J. Colmenar. 2017. Filling the Gondwanan gap: paleobiogeographic implications of new crinoids from the Castillejo and Fombuena formations (Middle and Upper Ordovician, Iberian Chains, Spain). Journal of Paleontology, v. 91, p. 715–734.

4. Wright, D.F., W.I. Ausich, S.R. Cole, T.W. Kammer, M.E. Peter, and E.C. Rhenberg. 2017. Phylogenetic taxonomy and classification of the Crinoidea (Phylum Echinodermata). Journal of Paleontology, v. 91, p. 829‒846.

3. Cole, S.R. and W.I. Ausich. 2015. Phylogenetic analysis of the Ordovician Diplobathrida (Subclass Camerata, Class Crinoidea): Implications for early camerate evolution. Cuadernos del Museo Geominero, v. 19: Progress in Echinoderm Palaeobiology, p. 41-44.

2. Ausich, W.I., T.W. Kammer, D.F. Wright, S.R. Cole, M.E. Peter, and E.C. Rhenberg. 2015. Toward a phylogenetic classification of the Crinoidea (Echinodermata). Cuadernos del Museo Geominero, no. 19: Progress in Echinoderm Palaeobiology, p. 29-32.

1. Cole, S.R., J. Haynes, P. Lucas, and R. Lambert. 2015. Faunal and sedimentological analysis of a latest Silurian stromatoporoid biostrome from the central Appalachian Basin. Facies, v. 61, no. 3, 16 p.